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The definitions and contents of the five species groups are as follows: leprus group: digital pads not or only slightly expanded, rounded in outline; first finger longer or shorter than second; snout acuminate or subacuminate, not rounded; outer metatarsal tubercle conical; digits lacking distinct lateral fringes, content: cystipnathoides, leprus and ruhrimactilatus. 8); venter white to cream; in life dorsum green with darker green spots, belly white; iris gold above, bronze below. — Cope's (1885) original description was not sufficiently clear to enable subsequent authors to recognize this species.
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HARVARD February 20, 1970 UHJVfi Wlbi TV A Taxonomic Revision of the Leptodactylid Frog Genus Syrrhophus Cope BY JOHN D. Lynch (1968) provided a definition of the pre- viously loosely-defined genus. presence or absence of vocal slits, and color pattern. Epirhexis never came into general usage ( Cope cited the name four times, but no one else has used it ) , whereas Syrrhophiis is well established in the zoological literature. Nuevo Leon: Salto Cola de Caballo, KU 92572; Huasteca Canyon, 15 km.
Taylor (1952) and Firschein (1954) limited the genus to several species of frogs occurring in Guatemala, Mexico, and Texas. Dennis, whose drawings great K' enhance the worth of this paper. Application of Baird's name Batrachijla longipes to the species of frog heretofore called Synluij^Jitis latodactyhis poses one serious problem. 13: Dorsal views of Syrrhophiis longipes illustrating geographic variation in pattern (left, TCWC 12179, xl.5; right, KU 92572, Xl.8); side of head (TCWC 10966, x6). However, since it seems almost certain that Batrac Jujla longipes and Syrrhophiis latodactyhis are conspecific, the former name should not be left as a nomen duhiiim. S Tacala, La Placita, 1850 m., FMNH 100266-68, 103244, UIMNH 13291, 13327. N Pablillo, EAL 1319; Sabinas Hidalgo, USNM 139728.
The distributions of some species have been extended, but otherwse the western complex of species remains unchanged since Duellman's review. Original combination Current combination campi, Syrrhophus cholorum, Syrrhophus lepriis cystigathoides, Ph yllobates dennisi, Syrrhophus gaigeae, Syrrho))lnis gutiilatus, Malacliylodes interorbitalis, Syrrhophus latodactylus, Syrrhophus leprus, Syrrhophus longipes, Batrachyla macrotympanum , Tomodactylus marnockii, Syrrhophus modcstus, Syrrhophus nehulosus, Syrrhoph us uiiocolimae, Syrrhophus ))al Udus, Syrrhophus modestus pctrophilus, Syrrhophus pipilans, Syrrhophus rubrintacidatus, Syrrhophus S77iithi, Syrrhophus teretistes, Syrrhophus verrticipes, Syrrhophus verruculatus, Phyllobates cystignathoides campi leprus cystignathoides cystignathoides dennisi new species guttilatus guttilatus interorbitalis longipes leprus longipes verrucipes marnockii modestus pipilans nehulosus nivocolimae pallidus guttilatus pipilans pipilans rubrimaculatus guttilatus teretistes verrucives Nomen duhium Leptodactylid Frog Genus Syrrhophus 5 with S. Thus, at present, nine species (one polytypic) are recognized on the eastern slopes and lowlands from central Texas to British Honduras. snout-\ent; vocal slits in males; digital tips greatly expanded, more than t^vice width of digit; first finger shorter than second; skin of dorsum shagreened to pustular, that of venter weakly to moderately areolate; toes webbed basally; dorsiun light brown to tan with brown \ermiculations; \enter wliite; diameter of tympanum 53.9 to 64.2 per cent that of eye in males, 50.6 to 58.7 per cent in females. First finger shorter than second; all fingers bearing truncate tips with pads, each pad having a terminal groo\e; fingers fringed; fingers three and four having dilated pads t\\o to three times width of digit; subarticular tubercles large, conical, rounded, simple; supernumerary tubercles nimierous on thenar surface, none on digits; three palmar tubercles, outer slightly smaller than largest super- numerary tubercles; row of tubercles on outer edge of forearm \ariable, weak to very distinct; tips of toes wider than digits, rounded to truncate at tips, each pad ha\ing terminal groo\e; toes ha\ing lateral fringes, bases of toes united by web, web not extending to basal subarticular tubercle; subarticular tubercles smaller than those of hand, round, conical, simple; supernumerary tubercles numerous on plantar surfaces, extending bet\veen metatarsal tubercles, present on toes between basal two subarticular tubercles in some specimens; outer metatarsal tubercle round, conical, one-half as large as ovoid, non-compressed inner metatarsal tubercle; tarsal tubercles or folds absent.
Dixon and Webb (1966) described an additional species from Jalisco, Mexico. marnockii) were recognized in Texas until Mil- stead, Mecham, and Mc Clintock (1950) synonymized S. Trivial name, original generic assignment, and author Current combination chalceus (Phyllobates) Peters, 1873 fesfae (Paludicola) Peracca, 1904 hylaeformis (Phyllobates) Cope, 1875 palmatus (Phyllobates) Werner, 1899 ridens (Phyllobates) Cope, 1866 simonsii (Paludicola) Boulenger, 1900 Eleutherodactylus chalceus Niceforonia festae Eleutherodactylus htjlaeformis Colostcthus palmatus Eleutherodactylus ridens Niceforonia simonsii Table 3 — Nominal Species of Syrrhophus (sensu strictu) and the Name Used Herein. Many scattered pustules on dorsum; flanks areolate; skin of venter areolate or not ( xariability may be due to difterences in preservation ) ; ventral disc distinct on chest and lower abdomen; inguinal gland present or not, when present varying from very large and distinct to poorly defined; axillary gland absent.
ZOOU University of K a n s a Sli^p^a oy Museum of Natural History ^v^AR /» I97Q Vol. These species are listed in Tables 1 and 2 with the name currently applied. Within the type-series, the pattern varies from \\'eakly to strongly \ermicnlate but is al\va\"s recognizable as vermiculate rather than spotted as in S. Therefore, Syrrhophus is used in this paper, even though Epirhexis has priority.
LYNCH University of Kansas Lawrence 1970 University of Kansas Publications, Museum of Nati^al History Editors of this number: Frank B. With the exception of Taylor (1952), who treated the Costa Rican species, none of these authors dealt with the present status of the nineteen species erroneously assigned to Sijrrhophus. Females of the two species can be separated by color pattern. It would serve only to confuse the Hterature to adhere strictly to the Law of Priority and replace Syrrhophus with Epirhexis.
I conclude that there are seven species (one poltypic) of Syrrhophus in eastern Mexico, Texas, and El Peten of Guatemala, and seven species (one polytypic) in western Mexico. Males of the two species can be readily separated by the relative sizes of the tympani, Leptodactylid Fhog Genus Syrrhophus 29 'W^ |U ^.:.;....,..
Dixon (1957) rede- fined the related genus Tomodactyhis and transferred T. It soon became evident that there were more names than species, that some previously regarded species were geographic \ ariants, and that the eastern and western groups (complexes here) were artificial divisions of the genus. latodachjlus." My stiidy indicates that the specimens from El Pachon represent a distinctive but allied species.
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